Mesosaur taxonomy reappraisal: are Stereosternum and Brazilosaurus valid taxa?
Resumen:
Mesosaurs are basal amniotes that lived at the beginning of the Permian or close to the Permo–Carboniferous boundary. Despite the several hundred specimens that have been found, including complete skeletons of adult and juvenile individuals, mesosaur taxonomy has been subjected to a high controversy over time. Currently, three monotypic genera, Mesosaurus tenuidens Gervais, Stereosternum tumidum Cope, and Brazilosaurus sanpauloensis Shikama & Ozaki are recognized, but identification of new specimens using the available diagnostic characters are arbitrary and influenced by high subjectivity. We performed anatomical and morphometric analyses to look for statistical support to these previously suggested basic diagnostic characters through an exhaustive anatomical revision of these characters and also of some new attributes discovered during the course of our study. We found a notable influence of taphonomic features in most of the diagnostic characters used to differentiate the three monotypic genera, including strong bias derived from the preservation of individuals in different ontogenetic stages, whose size and degree of ossification could have been controlled by particular environmental conditions that resulted in subtle polymorphisms of these and other few characters. Other features may even represent sexual dimorphism. After the detailed revision of the type specimens of the three currently acepted mesosaur taxa, for which we include here good-quality photographs, and considering the lack of statistical support for the most applied putative diagnostic features such as the different ratio found when comparing skull and cervical region lengths and the low or higher intensity of pachyosteosclerosis observed in dorsal ribs, which can be controlled by taphonomic and ecological conditions, we recognize Mesosaurus as the only mesosaurid taxon in the Paraná and Karoo basins, probably including dwarf individuals. Therefore, S. tumidum and B. sanpauloensis are suggested here as nomina dubia taking into account that the autapomorphies that supported these taxa cannot be confirmed to be absent in Mesosaurus.
Mesossauros são amniotas basais que viveram no início do Permiano ou próximo ao limite Permo–Carbonífero. A despeito das centenas de espécimes encontrados, incluindo esqueletos completos de indivíduos adultos e juvenis, a taxonomia dos mesossauros tem sido tema de muita controvérsia ao longo do tempo. Hoje, três gêneros monotípicos, Mesosaurus tenuidens Gervais, Stereosternum tumidum Cope, e Brazilosaurus sanpauloensis Shikama & Ozaki são reconhecidos, mas a identificação de novos espécimes utilizando os caracteres diagnósticos disponíveis é arbitrária e influenciada por alta subjetividade. Nós realizamos análises anatômica e morfométrica, tendo em vista obter um suporte estatístico para estes caracteres sugeridos como diagnósticos básicos por meio de uma exaustiva revisão anatômica destes caracteres, e de alguns novos caracteres descobertos durante o desenvolvimento de nosso estudo. Nós notamos uma notável influência de fatores tafonômicos na maioria dos caracteres diagnósticos utilizados para os três táxons monotípicos e incluindo uma forte preconcepção derivada da preservação dos indivíduos em diferentes estágios ontogenéticos, cujos tamanho e grau de ossificação poderiam ser controlados por condições ambientais que resultaram em sutis polimorfismos destes poucos caracteres. Outros aspectos podem mesmo representar dimorfismo sexual. Considerando a falta de suporte estatístico para a maioria dos putativos caracteres diagnósticos, tais como as diferentes relações entre comprimento do crânio e da coluna cervical e a pressão das condições tafonômicas e ecológicas, que podem ter controlado um menor ou maior grau de paquiosteoesclerose nas costelas dorsais, nós reconhecemos Mesosaurus como o único táxon de mesosaurídeo nas bacias do Paraná e Karoo, provavelmente incluindo populações anãs, e sugerimos que S. tumidum e B. sanpauloensis devem ser considerados como nomen dubium, levando em conta que as autapomorfias que suportam estes táxons não podem ser confirmadas como não estando presentes em Mesosaurus.
2021 | |
Mesosaurus Morphometrics Taxonomy ?Early Permian Gondwana |
|
Inglés | |
Universidad de la República | |
COLIBRI | |
https://hdl.handle.net/20.500.12008/41441 | |
Acceso abierto | |
Licencia Creative Commons Atribución (CC - By 4.0) |
_version_ | 1807522802516885504 |
---|---|
author | Piñeiro Martínez, Graciela H. |
author2 | Ferigolo, Jorge Mones, Alvaro Núñez Demarco, Pablo Andrés |
author2_role | author author author |
author_facet | Piñeiro Martínez, Graciela H. Ferigolo, Jorge Mones, Alvaro Núñez Demarco, Pablo Andrés |
author_role | author |
bitstream.checksum.fl_str_mv | 6429389a7df7277b72b7924fdc7d47a9 a0ebbeafb9d2ec7cbb19d7137ebc392c 9b2f65dcc322aef148d393163faed91b 71ed42ef0a0b648670f707320be37b90 dd40322adec80ba2ec438fbabdfc9f35 |
bitstream.checksumAlgorithm.fl_str_mv | MD5 MD5 MD5 MD5 MD5 |
bitstream.url.fl_str_mv | http://localhost:8080/xmlui/bitstream/20.500.12008/41441/5/license.txt http://localhost:8080/xmlui/bitstream/20.500.12008/41441/2/license_url http://localhost:8080/xmlui/bitstream/20.500.12008/41441/3/license_text http://localhost:8080/xmlui/bitstream/20.500.12008/41441/4/license_rdf http://localhost:8080/xmlui/bitstream/20.500.12008/41441/1/104072rbp2021304.pdf |
collection | COLIBRI |
dc.contributor.filiacion.none.fl_str_mv | Piñeiro Martínez Graciela H., Universidad de la República (Uruguay). Facultad de Ciencias. Instituto de Ciencias Geológicas. Ferigolo Jorge Mones Alvaro, MNHN (Uruguay) Núñez Demarco Pablo Andrés, Universidad de la República (Uruguay). Facultad de Ciencias. Instituto de Ciencias Geológicas. |
dc.creator.none.fl_str_mv | Piñeiro Martínez, Graciela H. Ferigolo, Jorge Mones, Alvaro Núñez Demarco, Pablo Andrés |
dc.date.accessioned.none.fl_str_mv | 2023-11-22T17:48:57Z |
dc.date.available.none.fl_str_mv | 2023-11-22T17:48:57Z |
dc.date.issued.none.fl_str_mv | 2021 |
dc.description.abstract.none.fl_txt_mv | Mesosaurs are basal amniotes that lived at the beginning of the Permian or close to the Permo–Carboniferous boundary. Despite the several hundred specimens that have been found, including complete skeletons of adult and juvenile individuals, mesosaur taxonomy has been subjected to a high controversy over time. Currently, three monotypic genera, Mesosaurus tenuidens Gervais, Stereosternum tumidum Cope, and Brazilosaurus sanpauloensis Shikama & Ozaki are recognized, but identification of new specimens using the available diagnostic characters are arbitrary and influenced by high subjectivity. We performed anatomical and morphometric analyses to look for statistical support to these previously suggested basic diagnostic characters through an exhaustive anatomical revision of these characters and also of some new attributes discovered during the course of our study. We found a notable influence of taphonomic features in most of the diagnostic characters used to differentiate the three monotypic genera, including strong bias derived from the preservation of individuals in different ontogenetic stages, whose size and degree of ossification could have been controlled by particular environmental conditions that resulted in subtle polymorphisms of these and other few characters. Other features may even represent sexual dimorphism. After the detailed revision of the type specimens of the three currently acepted mesosaur taxa, for which we include here good-quality photographs, and considering the lack of statistical support for the most applied putative diagnostic features such as the different ratio found when comparing skull and cervical region lengths and the low or higher intensity of pachyosteosclerosis observed in dorsal ribs, which can be controlled by taphonomic and ecological conditions, we recognize Mesosaurus as the only mesosaurid taxon in the Paraná and Karoo basins, probably including dwarf individuals. Therefore, S. tumidum and B. sanpauloensis are suggested here as nomina dubia taking into account that the autapomorphies that supported these taxa cannot be confirmed to be absent in Mesosaurus. Mesossauros são amniotas basais que viveram no início do Permiano ou próximo ao limite Permo–Carbonífero. A despeito das centenas de espécimes encontrados, incluindo esqueletos completos de indivíduos adultos e juvenis, a taxonomia dos mesossauros tem sido tema de muita controvérsia ao longo do tempo. Hoje, três gêneros monotípicos, Mesosaurus tenuidens Gervais, Stereosternum tumidum Cope, e Brazilosaurus sanpauloensis Shikama & Ozaki são reconhecidos, mas a identificação de novos espécimes utilizando os caracteres diagnósticos disponíveis é arbitrária e influenciada por alta subjetividade. Nós realizamos análises anatômica e morfométrica, tendo em vista obter um suporte estatístico para estes caracteres sugeridos como diagnósticos básicos por meio de uma exaustiva revisão anatômica destes caracteres, e de alguns novos caracteres descobertos durante o desenvolvimento de nosso estudo. Nós notamos uma notável influência de fatores tafonômicos na maioria dos caracteres diagnósticos utilizados para os três táxons monotípicos e incluindo uma forte preconcepção derivada da preservação dos indivíduos em diferentes estágios ontogenéticos, cujos tamanho e grau de ossificação poderiam ser controlados por condições ambientais que resultaram em sutis polimorfismos destes poucos caracteres. Outros aspectos podem mesmo representar dimorfismo sexual. Considerando a falta de suporte estatístico para a maioria dos putativos caracteres diagnósticos, tais como as diferentes relações entre comprimento do crânio e da coluna cervical e a pressão das condições tafonômicas e ecológicas, que podem ter controlado um menor ou maior grau de paquiosteoesclerose nas costelas dorsais, nós reconhecemos Mesosaurus como o único táxon de mesosaurídeo nas bacias do Paraná e Karoo, provavelmente incluindo populações anãs, e sugerimos que S. tumidum e B. sanpauloensis devem ser considerados como nomen dubium, levando em conta que as autapomorfias que suportam estes táxons não podem ser confirmadas como não estando presentes em Mesosaurus. |
dc.format.extent.es.fl_str_mv | 31 h. |
dc.format.mimetype.es.fl_str_mv | application/pdf |
dc.identifier.citation.es.fl_str_mv | Piñeiro Martínez, G, Ferigolo, J, Mones, A [y otro autor]. "Mesosaur taxonomy reappraisal: are Stereosternum and Brazilosaurus valid taxa?". Revista Brasileira de Paleontologia. [en línea] 2021, 24(3): 205-235. 31 h. DOI: 10.4072/rbp.2021.3.04. |
dc.identifier.doi.none.fl_str_mv | 10.4072/rbp.2021.3.04 |
dc.identifier.issn.none.fl_str_mv | 2236-1715 |
dc.identifier.uri.none.fl_str_mv | https://hdl.handle.net/20.500.12008/41441 |
dc.language.iso.none.fl_str_mv | en eng |
dc.publisher.es.fl_str_mv | Brazilian Society of Paleontology |
dc.relation.ispartof.es.fl_str_mv | Revista Brasileira de Paleontologia, 2021, 24(3): 205-235. |
dc.rights.license.none.fl_str_mv | Licencia Creative Commons Atribución (CC - By 4.0) |
dc.rights.none.fl_str_mv | info:eu-repo/semantics/openAccess |
dc.source.none.fl_str_mv | reponame:COLIBRI instname:Universidad de la República instacron:Universidad de la República |
dc.subject.es.fl_str_mv | Mesosaurus Morphometrics Taxonomy ?Early Permian Gondwana |
dc.title.none.fl_str_mv | Mesosaur taxonomy reappraisal: are Stereosternum and Brazilosaurus valid taxa? |
dc.type.es.fl_str_mv | Artículo |
dc.type.none.fl_str_mv | info:eu-repo/semantics/article |
dc.type.version.none.fl_str_mv | info:eu-repo/semantics/publishedVersion |
description | Mesosaurs are basal amniotes that lived at the beginning of the Permian or close to the Permo–Carboniferous boundary. Despite the several hundred specimens that have been found, including complete skeletons of adult and juvenile individuals, mesosaur taxonomy has been subjected to a high controversy over time. Currently, three monotypic genera, Mesosaurus tenuidens Gervais, Stereosternum tumidum Cope, and Brazilosaurus sanpauloensis Shikama & Ozaki are recognized, but identification of new specimens using the available diagnostic characters are arbitrary and influenced by high subjectivity. We performed anatomical and morphometric analyses to look for statistical support to these previously suggested basic diagnostic characters through an exhaustive anatomical revision of these characters and also of some new attributes discovered during the course of our study. We found a notable influence of taphonomic features in most of the diagnostic characters used to differentiate the three monotypic genera, including strong bias derived from the preservation of individuals in different ontogenetic stages, whose size and degree of ossification could have been controlled by particular environmental conditions that resulted in subtle polymorphisms of these and other few characters. Other features may even represent sexual dimorphism. After the detailed revision of the type specimens of the three currently acepted mesosaur taxa, for which we include here good-quality photographs, and considering the lack of statistical support for the most applied putative diagnostic features such as the different ratio found when comparing skull and cervical region lengths and the low or higher intensity of pachyosteosclerosis observed in dorsal ribs, which can be controlled by taphonomic and ecological conditions, we recognize Mesosaurus as the only mesosaurid taxon in the Paraná and Karoo basins, probably including dwarf individuals. Therefore, S. tumidum and B. sanpauloensis are suggested here as nomina dubia taking into account that the autapomorphies that supported these taxa cannot be confirmed to be absent in Mesosaurus. |
eu_rights_str_mv | openAccess |
format | article |
id | COLIBRI_7590bea7e31465acb08469d7039943a6 |
identifier_str_mv | Piñeiro Martínez, G, Ferigolo, J, Mones, A [y otro autor]. "Mesosaur taxonomy reappraisal: are Stereosternum and Brazilosaurus valid taxa?". Revista Brasileira de Paleontologia. [en línea] 2021, 24(3): 205-235. 31 h. DOI: 10.4072/rbp.2021.3.04. 2236-1715 10.4072/rbp.2021.3.04 |
instacron_str | Universidad de la República |
institution | Universidad de la República |
instname_str | Universidad de la República |
language | eng |
language_invalid_str_mv | en |
network_acronym_str | COLIBRI |
network_name_str | COLIBRI |
oai_identifier_str | oai:colibri.udelar.edu.uy:20.500.12008/41441 |
publishDate | 2021 |
reponame_str | COLIBRI |
repository.mail.fl_str_mv | mabel.seroubian@seciu.edu.uy |
repository.name.fl_str_mv | COLIBRI - Universidad de la República |
repository_id_str | 4771 |
rights_invalid_str_mv | Licencia Creative Commons Atribución (CC - By 4.0) |
spelling | Piñeiro Martínez Graciela H., Universidad de la República (Uruguay). Facultad de Ciencias. Instituto de Ciencias Geológicas.Ferigolo JorgeMones Alvaro, MNHN (Uruguay)Núñez Demarco Pablo Andrés, Universidad de la República (Uruguay). Facultad de Ciencias. Instituto de Ciencias Geológicas.2023-11-22T17:48:57Z2023-11-22T17:48:57Z2021Piñeiro Martínez, G, Ferigolo, J, Mones, A [y otro autor]. "Mesosaur taxonomy reappraisal: are Stereosternum and Brazilosaurus valid taxa?". Revista Brasileira de Paleontologia. [en línea] 2021, 24(3): 205-235. 31 h. DOI: 10.4072/rbp.2021.3.04.2236-1715https://hdl.handle.net/20.500.12008/4144110.4072/rbp.2021.3.04Mesosaurs are basal amniotes that lived at the beginning of the Permian or close to the Permo–Carboniferous boundary. Despite the several hundred specimens that have been found, including complete skeletons of adult and juvenile individuals, mesosaur taxonomy has been subjected to a high controversy over time. Currently, three monotypic genera, Mesosaurus tenuidens Gervais, Stereosternum tumidum Cope, and Brazilosaurus sanpauloensis Shikama & Ozaki are recognized, but identification of new specimens using the available diagnostic characters are arbitrary and influenced by high subjectivity. We performed anatomical and morphometric analyses to look for statistical support to these previously suggested basic diagnostic characters through an exhaustive anatomical revision of these characters and also of some new attributes discovered during the course of our study. We found a notable influence of taphonomic features in most of the diagnostic characters used to differentiate the three monotypic genera, including strong bias derived from the preservation of individuals in different ontogenetic stages, whose size and degree of ossification could have been controlled by particular environmental conditions that resulted in subtle polymorphisms of these and other few characters. Other features may even represent sexual dimorphism. After the detailed revision of the type specimens of the three currently acepted mesosaur taxa, for which we include here good-quality photographs, and considering the lack of statistical support for the most applied putative diagnostic features such as the different ratio found when comparing skull and cervical region lengths and the low or higher intensity of pachyosteosclerosis observed in dorsal ribs, which can be controlled by taphonomic and ecological conditions, we recognize Mesosaurus as the only mesosaurid taxon in the Paraná and Karoo basins, probably including dwarf individuals. Therefore, S. tumidum and B. sanpauloensis are suggested here as nomina dubia taking into account that the autapomorphies that supported these taxa cannot be confirmed to be absent in Mesosaurus.Mesossauros são amniotas basais que viveram no início do Permiano ou próximo ao limite Permo–Carbonífero. A despeito das centenas de espécimes encontrados, incluindo esqueletos completos de indivíduos adultos e juvenis, a taxonomia dos mesossauros tem sido tema de muita controvérsia ao longo do tempo. Hoje, três gêneros monotípicos, Mesosaurus tenuidens Gervais, Stereosternum tumidum Cope, e Brazilosaurus sanpauloensis Shikama & Ozaki são reconhecidos, mas a identificação de novos espécimes utilizando os caracteres diagnósticos disponíveis é arbitrária e influenciada por alta subjetividade. Nós realizamos análises anatômica e morfométrica, tendo em vista obter um suporte estatístico para estes caracteres sugeridos como diagnósticos básicos por meio de uma exaustiva revisão anatômica destes caracteres, e de alguns novos caracteres descobertos durante o desenvolvimento de nosso estudo. Nós notamos uma notável influência de fatores tafonômicos na maioria dos caracteres diagnósticos utilizados para os três táxons monotípicos e incluindo uma forte preconcepção derivada da preservação dos indivíduos em diferentes estágios ontogenéticos, cujos tamanho e grau de ossificação poderiam ser controlados por condições ambientais que resultaram em sutis polimorfismos destes poucos caracteres. Outros aspectos podem mesmo representar dimorfismo sexual. Considerando a falta de suporte estatístico para a maioria dos putativos caracteres diagnósticos, tais como as diferentes relações entre comprimento do crânio e da coluna cervical e a pressão das condições tafonômicas e ecológicas, que podem ter controlado um menor ou maior grau de paquiosteoesclerose nas costelas dorsais, nós reconhecemos Mesosaurus como o único táxon de mesosaurídeo nas bacias do Paraná e Karoo, provavelmente incluindo populações anãs, e sugerimos que S. tumidum e B. sanpauloensis devem ser considerados como nomen dubium, levando em conta que as autapomorfias que suportam estes táxons não podem ser confirmadas como não estando presentes em Mesosaurus.Submitted by Parodi Mónica (mparodi@fcien.edu.uy) on 2023-11-21T17:11:38Z No. of bitstreams: 2 license_rdf: 24251 bytes, checksum: 71ed42ef0a0b648670f707320be37b90 (MD5) 104072rbp2021304.pdf: 3403134 bytes, checksum: dd40322adec80ba2ec438fbabdfc9f35 (MD5)Approved for entry into archive by Faget Cecilia (lfaget@fcien.edu.uy) on 2023-11-21T17:33:42Z (GMT) No. of bitstreams: 2 license_rdf: 24251 bytes, checksum: 71ed42ef0a0b648670f707320be37b90 (MD5) 104072rbp2021304.pdf: 3403134 bytes, checksum: dd40322adec80ba2ec438fbabdfc9f35 (MD5)Made available in DSpace by Luna Fabiana (fabiana.luna@seciu.edu.uy) on 2023-11-22T17:48:57Z (GMT). No. of bitstreams: 2 license_rdf: 24251 bytes, checksum: 71ed42ef0a0b648670f707320be37b90 (MD5) 104072rbp2021304.pdf: 3403134 bytes, checksum: dd40322adec80ba2ec438fbabdfc9f35 (MD5) Previous issue date: 202131 h.application/pdfenengBrazilian Society of PaleontologyRevista Brasileira de Paleontologia, 2021, 24(3): 205-235.Las obras depositadas en el Repositorio se rigen por la Ordenanza de los Derechos de la Propiedad Intelectual de la Universidad de la República.(Res. Nº 91 de C.D.C. de 8/III/1994 – D.O. 7/IV/1994) y por la Ordenanza del Repositorio Abierto de la Universidad de la República (Res. Nº 16 de C.D.C. de 07/10/2014)info:eu-repo/semantics/openAccessLicencia Creative Commons Atribución (CC - By 4.0)MesosaurusMorphometricsTaxonomy?Early PermianGondwanaMesosaur taxonomy reappraisal: are Stereosternum and Brazilosaurus valid taxa?Artículoinfo:eu-repo/semantics/articleinfo:eu-repo/semantics/publishedVersionreponame:COLIBRIinstname:Universidad de la Repúblicainstacron:Universidad de la RepúblicaPiñeiro Martínez, Graciela H.Ferigolo, JorgeMones, AlvaroNúñez Demarco, Pablo AndrésLICENSElicense.txtlicense.txttext/plain; charset=utf-84267http://localhost:8080/xmlui/bitstream/20.500.12008/41441/5/license.txt6429389a7df7277b72b7924fdc7d47a9MD55CC-LICENSElicense_urllicense_urltext/plain; charset=utf-844http://localhost:8080/xmlui/bitstream/20.500.12008/41441/2/license_urla0ebbeafb9d2ec7cbb19d7137ebc392cMD52license_textlicense_texttext/html; charset=utf-813786http://localhost:8080/xmlui/bitstream/20.500.12008/41441/3/license_text9b2f65dcc322aef148d393163faed91bMD53license_rdflicense_rdfapplication/rdf+xml; charset=utf-824251http://localhost:8080/xmlui/bitstream/20.500.12008/41441/4/license_rdf71ed42ef0a0b648670f707320be37b90MD54ORIGINAL104072rbp2021304.pdf104072rbp2021304.pdfapplication/pdf3403134http://localhost:8080/xmlui/bitstream/20.500.12008/41441/1/104072rbp2021304.pdfdd40322adec80ba2ec438fbabdfc9f35MD5120.500.12008/414412023-11-22 14:48:57.63oai:colibri.udelar.edu.uy:20.500.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Universidadhttps://udelar.edu.uy/https://www.colibri.udelar.edu.uy/oai/requestmabel.seroubian@seciu.edu.uyUruguayopendoar:47712024-07-25T14:29:12.494808COLIBRI - Universidad de la Repúblicafalse |
spellingShingle | Mesosaur taxonomy reappraisal: are Stereosternum and Brazilosaurus valid taxa? Piñeiro Martínez, Graciela H. Mesosaurus Morphometrics Taxonomy ?Early Permian Gondwana |
status_str | publishedVersion |
title | Mesosaur taxonomy reappraisal: are Stereosternum and Brazilosaurus valid taxa? |
title_full | Mesosaur taxonomy reappraisal: are Stereosternum and Brazilosaurus valid taxa? |
title_fullStr | Mesosaur taxonomy reappraisal: are Stereosternum and Brazilosaurus valid taxa? |
title_full_unstemmed | Mesosaur taxonomy reappraisal: are Stereosternum and Brazilosaurus valid taxa? |
title_short | Mesosaur taxonomy reappraisal: are Stereosternum and Brazilosaurus valid taxa? |
title_sort | Mesosaur taxonomy reappraisal: are Stereosternum and Brazilosaurus valid taxa? |
topic | Mesosaurus Morphometrics Taxonomy ?Early Permian Gondwana |
url | https://hdl.handle.net/20.500.12008/41441 |