Enigmatic wood and first evidence of tetrapods in the Yaguarí Formation (Middle-Late Permian), Uruguay

Madera enigmática y primera evidencia de tetrápodos en Formación Yaguarí (Pérmico Medio-Superior), Uruguay

Madeira enigmática e primeira evidência de tetrápodes na Formação Yaguarí (Permiano Médio-Superior), Uruguai

Piñeiro, Graciela Helena - Marchetti, Lorenzo - Marmol, Sebastián - Celio, Antonella - Xavier, Pedro Luis - Francia, Martin - Schultz, Cesar Leandro

Editor(es): Pérez-Bidegain, Mario - Gaucher, Claudio

Resumen:

This article describes an intriguing fossil material recently found in the fine-grained sandstone levels from the upper section of the Yaguarí Formation of Uruguay. The nature of this specimen and its relationships were, initially, not easy to determine with confidence although it was subjected to several analyses, such as morphological and comparative examination complemented with preparation of thin sections and microanatomical and chemical studies under SEM. However, the anatomical structure shown in the performed transversal thin sections allowed solving the issue on the different animal or plant interpretations of the fossil specimen, which was finally discerned in favor of the latter. Therefore, after a detailed comparative study of the thin sections, we concluded that the specimen is a wood fragment related to the Dadoxylon-Araucarioxylon complex. Moreover, in order to provide a better biostratigraphic calibration for these deposits until now devoid of fossils, we include a preliminary description of recently found fossil footprints, which represent the first evidence of the presence of tetrapods in levels of the Yaguarí Formation (sensu Elizalde et al., 1970), below the conglomerates that have yielded the Colonia Orozco Fauna. The footprints were preliminary assigned to the ichnotaxa cf. Karoopes isp, cf. Capitosauroides isp., and cf. Pachypesisp., suggesting the presence of gorgonopsids and therocephalian therapsids and pareiasauromorphs, respectively. The Middle-Late Permian age recently suggested for the Yaguarí Formation based on paleomagnetic and radioisotopic studies is here supported by the described fossils, which thus constitute preliminary but nonetheless relevant discoveries for deposits whose fossiliferous potential has yet to be explored.


Este artículo describe un intrigante fósil recientemente descubierto en niveles de areniscas finas a medias de la sección superior de la Formación Yaguarí de Uruguay. La naturaleza y las afinidades de este espécimen no fueron inicialmente fáciles de determinar, a pesar de ser sometido a numerosos estudios, incluido un análisis anatómico-comparativo complementado con la preparación de secciones delgadas y análisis de su microestructura bajo Microscopio Electrónico de Barrido. La microestructura mostrada por las láminas delgadas en corte transversal permitió finalmente resolver el problema sobre discernir entre una naturaleza animal o vegetal del espécimen, siendo la última la aceptada. El detallado estudio comparativo de las secciones delgadas reveló que el espécimen es un fragmento de madera relacionado al complejo Dadoxylon-Araucarioxylon. Asimismo, para proveer una mejor calibración temporal de los depósitos de la sección superior de la Formación Yaguarí (Miembro Villa Viñoles), también se incluye en este trabajo una descripción preliminar de huellas de tetrápodos recientemente halladas en esta unidad, las cuales representan la primera evidencia de la presencia de tetrápodos en niveles de la Formación Yaguarí (sensu Elizalde et al., 1970) subyacentes a los conglomerados portadores de la Fauna de Colonia Orozco. Las huellas fueron asignadas a los icnotaxa cf. Karoopes isp., cf. Capitosauroides isp. y cf. Pachypes isp., sugiriendo la presencia de terápsidos gorgonópsidos y therocefalidos y de pareiasauromorfos, respectivamente. La edad Pérmico Medio a Tardío recientemente sugerida para la Formación Yaguarí, basado en estudios de paleomagnetismo sustentados por dataciones radioisotópicas, es congruente con la aportada por los fósiles aquí descriptos, que, aunque fragmentarios, constituyen descubrimientos preliminarmente relevantes para depósitos cuyo potencial fosilífero no ha sido aún explorado.


Este artigo descreve um intrigante material fóssil achado recentemente nos níveis de arenito fino da seção superior da Formação Yaguarí no Uruguai. A natureza deste espécime e suas relações não foram, em um primeiro momento, facilmente determináveis, apesar do mesmo ter sido submetido a várias análises. Estas incluíram avaliações preliminares comparativas e de morfologia, complementadas com preparação de lâminas delgadas, e estudos microanatômicos e químicos em MEV. A estrutura anatômica observada nas seções delgadas transversais permitiu solucionar as controversas interpretações do espécime fóssil, animal ou planta, o qual foi finalmente discernido em favor da última. Por enquanto, depois do estudo detalhado das lâminas delgadas o espécime foi relacionado a o complexo Dadoxylon-Araucarioxylon. Além disso, para permitir uma melhor calibração bioestratigráfica para estes depósitos até então desprovidos de fósseis, incluímos uma descrição preliminar de pegadas fósseis recém-encontradas, as quais são a primeira evidência a presença de tetrápodes em níveis da Formação Yaguarí (sensu Elizalde et al., 1970), abaixo dos conglomerados que forneceram a Fauna de Colonia Orozco. As pegadas foram assignadas aos ichnotaxa cf. Karoopes isp., cf. Capitosauroides isp. e cf. Pachypes isp., sugerindo a presença de terápsidos gorgonopsídeos e terocefalídeos e de pareiasauromorfos, respectivamente. Uma idade Meso-Neopermiano recentemente sugerida para a Formação Yaguarí baseada em estudos paleomagnéticos e radiométricos é aqui suportada pelos fósseis descritos, os quais, apesar de serem descobertas preliminares, fornecem dados relevantes para depósitos cujo potencial fossilífero ainda está para ser explorado.


Detalles Bibliográficos
2022
Capitosauroides and Pachypes-Like Tracks
Capitosauroides e Pachypes
Capitosauroides y Pachypes
Dadoxylon-Like Wood
Estratigrafia da Formação Yaguarí
Estratigrafía de Formación Yaguarí
Guadalupian-Lopingian Boundary
Huellas Karoopes
Karoopes
Limite Guadalupiano-Lopingiano
Límite Guadalupiano-Lopingiano
Madeiras Dadoxylon
Maderas Dadoxylon
Paleoambientes
Paleoenvironments
Pegadas Karoopes
Uruguai
Uruguay
Yaguarí Formation Stratigraphy
Inglés
Universidad de la República
COLIBRI
https://hdl.handle.net/20.500.12008/43673
Acceso abierto
Licencia Creative Commons Atribución (CC - By 4.0)
_version_ 1807523264525762560
author Piñeiro, Graciela Helena
author2 Marchetti, Lorenzo
Marmol, Sebastián
Celio, Antonella
Xavier, Pedro Luis
Francia, Martin
Schultz, Cesar Leandro
author2_role author
author
author
author
author
author
author_facet Piñeiro, Graciela Helena
Marchetti, Lorenzo
Marmol, Sebastián
Celio, Antonella
Xavier, Pedro Luis
Francia, Martin
Schultz, Cesar Leandro
author_role author
bitstream.checksum.fl_str_mv 6429389a7df7277b72b7924fdc7d47a9
a0ebbeafb9d2ec7cbb19d7137ebc392c
3fd1ebed2dc74f31b189736d0c5b4212
71ed42ef0a0b648670f707320be37b90
6b62e0880b09cc79ac1c26fbd0949760
bitstream.checksumAlgorithm.fl_str_mv MD5
MD5
MD5
MD5
MD5
bitstream.url.fl_str_mv http://localhost:8080/xmlui/bitstream/20.500.12008/43673/5/license.txt
http://localhost:8080/xmlui/bitstream/20.500.12008/43673/2/license_url
http://localhost:8080/xmlui/bitstream/20.500.12008/43673/3/license_text
http://localhost:8080/xmlui/bitstream/20.500.12008/43673/4/license_rdf
http://localhost:8080/xmlui/bitstream/20.500.12008/43673/1/2730-5066--504.pdf
collection COLIBRI
dc.contributor.filiacion.none.fl_str_mv Piñeiro Graciela Helena, Universidad de la República, Facultad de Ciencias, Departamento de Paleontología, Montevideo, Uruguay https://orcid.org/0000-0002-5872-9721
Marchetti Lorenzo, Museum für Naturkunde Berlin, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Berlin, Germany https://orcid.org/0000-0002-1047-7887
Marmol Sebastián, Universidad de la República, Facultad de Agronomía, Departamento de suelos y aguas, Montevideo, Uruguay https://orcid.org/0000-0001-7059-2724
Celio Antonella, Universidad de la República, Facultad de Agronomía, Departamento de suelos y aguas, Montevideo, Uruguay https://orcid.org/0000-0002-1425-0217
Xavier Pedro Luis, Universidade Federal do Rio Grande do Sul, Programa de Pós-Graduação em Geociências, Porto Alegre, Brasil https://orcid.org/0000-0001-9090-5361
Francia Martin, Universidad de la República, Facultad de Agronomía, Departamento de suelos y aguas, Montevideo, Uruguay https://orcid.org/0000-0001-6107-2815
Schultz Cesar Leandro, Universidade Federal do Rio Grande do Sul, Departamento de Estratigrafia e Paleontologia, Porto Alegre, Brasil https://orcid.org/0000-0001-7121-0409
dc.creator.editor.none.fl_str_mv Pérez-Bidegain, Mario
Gaucher, Claudio
dc.creator.none.fl_str_mv Piñeiro, Graciela Helena
Marchetti, Lorenzo
Marmol, Sebastián
Celio, Antonella
Xavier, Pedro Luis
Francia, Martin
Schultz, Cesar Leandro
dc.date.accessioned.none.fl_str_mv 2024-04-29T14:56:21Z
dc.date.available.none.fl_str_mv 2024-04-29T14:56:21Z
dc.date.issued.none.fl_str_mv 2022
dc.description.abstract.none.fl_txt_mv This article describes an intriguing fossil material recently found in the fine-grained sandstone levels from the upper section of the Yaguarí Formation of Uruguay. The nature of this specimen and its relationships were, initially, not easy to determine with confidence although it was subjected to several analyses, such as morphological and comparative examination complemented with preparation of thin sections and microanatomical and chemical studies under SEM. However, the anatomical structure shown in the performed transversal thin sections allowed solving the issue on the different animal or plant interpretations of the fossil specimen, which was finally discerned in favor of the latter. Therefore, after a detailed comparative study of the thin sections, we concluded that the specimen is a wood fragment related to the Dadoxylon-Araucarioxylon complex. Moreover, in order to provide a better biostratigraphic calibration for these deposits until now devoid of fossils, we include a preliminary description of recently found fossil footprints, which represent the first evidence of the presence of tetrapods in levels of the Yaguarí Formation (sensu Elizalde et al., 1970), below the conglomerates that have yielded the Colonia Orozco Fauna. The footprints were preliminary assigned to the ichnotaxa cf. Karoopes isp, cf. Capitosauroides isp., and cf. Pachypesisp., suggesting the presence of gorgonopsids and therocephalian therapsids and pareiasauromorphs, respectively. The Middle-Late Permian age recently suggested for the Yaguarí Formation based on paleomagnetic and radioisotopic studies is here supported by the described fossils, which thus constitute preliminary but nonetheless relevant discoveries for deposits whose fossiliferous potential has yet to be explored.
Este artículo describe un intrigante fósil recientemente descubierto en niveles de areniscas finas a medias de la sección superior de la Formación Yaguarí de Uruguay. La naturaleza y las afinidades de este espécimen no fueron inicialmente fáciles de determinar, a pesar de ser sometido a numerosos estudios, incluido un análisis anatómico-comparativo complementado con la preparación de secciones delgadas y análisis de su microestructura bajo Microscopio Electrónico de Barrido. La microestructura mostrada por las láminas delgadas en corte transversal permitió finalmente resolver el problema sobre discernir entre una naturaleza animal o vegetal del espécimen, siendo la última la aceptada. El detallado estudio comparativo de las secciones delgadas reveló que el espécimen es un fragmento de madera relacionado al complejo Dadoxylon-Araucarioxylon. Asimismo, para proveer una mejor calibración temporal de los depósitos de la sección superior de la Formación Yaguarí (Miembro Villa Viñoles), también se incluye en este trabajo una descripción preliminar de huellas de tetrápodos recientemente halladas en esta unidad, las cuales representan la primera evidencia de la presencia de tetrápodos en niveles de la Formación Yaguarí (sensu Elizalde et al., 1970) subyacentes a los conglomerados portadores de la Fauna de Colonia Orozco. Las huellas fueron asignadas a los icnotaxa cf. Karoopes isp., cf. Capitosauroides isp. y cf. Pachypes isp., sugiriendo la presencia de terápsidos gorgonópsidos y therocefalidos y de pareiasauromorfos, respectivamente. La edad Pérmico Medio a Tardío recientemente sugerida para la Formación Yaguarí, basado en estudios de paleomagnetismo sustentados por dataciones radioisotópicas, es congruente con la aportada por los fósiles aquí descriptos, que, aunque fragmentarios, constituyen descubrimientos preliminarmente relevantes para depósitos cuyo potencial fosilífero no ha sido aún explorado.
Este artigo descreve um intrigante material fóssil achado recentemente nos níveis de arenito fino da seção superior da Formação Yaguarí no Uruguai. A natureza deste espécime e suas relações não foram, em um primeiro momento, facilmente determináveis, apesar do mesmo ter sido submetido a várias análises. Estas incluíram avaliações preliminares comparativas e de morfologia, complementadas com preparação de lâminas delgadas, e estudos microanatômicos e químicos em MEV. A estrutura anatômica observada nas seções delgadas transversais permitiu solucionar as controversas interpretações do espécime fóssil, animal ou planta, o qual foi finalmente discernido em favor da última. Por enquanto, depois do estudo detalhado das lâminas delgadas o espécime foi relacionado a o complexo Dadoxylon-Araucarioxylon. Além disso, para permitir uma melhor calibração bioestratigráfica para estes depósitos até então desprovidos de fósseis, incluímos uma descrição preliminar de pegadas fósseis recém-encontradas, as quais são a primeira evidência a presença de tetrápodes em níveis da Formação Yaguarí (sensu Elizalde et al., 1970), abaixo dos conglomerados que forneceram a Fauna de Colonia Orozco. As pegadas foram assignadas aos ichnotaxa cf. Karoopes isp., cf. Capitosauroides isp. e cf. Pachypes isp., sugerindo a presença de terápsidos gorgonopsídeos e terocefalídeos e de pareiasauromorfos, respectivamente. Uma idade Meso-Neopermiano recentemente sugerida para a Formação Yaguarí baseada em estudos paleomagnéticos e radiométricos é aqui suportada pelos fósseis descritos, os quais, apesar de serem descobertas preliminares, fornecem dados relevantes para depósitos cujo potencial fossilífero ainda está para ser explorado.
dc.format.extent.es.fl_str_mv e504
dc.format.mimetype.es.fl_str_mv application/pdf
dc.identifier.citation.es.fl_str_mv Piñeiro, G. H., Marchetti, L., Marmol, S., y otros. Enigmatic wood and first evidence of tetrapods in the Yaguarí Formation (Middle-Late Permian), Uruguay. Agrociencia Uruguay, 26(NE1), e504. https://doi.org/10.31285/AGRO.26.504
dc.identifier.doi.none.fl_str_mv 10.31285/AGRO.26.504
dc.identifier.issn.none.fl_str_mv 2730-5066
dc.identifier.uri.none.fl_str_mv https://hdl.handle.net/20.500.12008/43673
dc.language.iso.none.fl_str_mv en
eng
dc.publisher.es.fl_str_mv Facultad de Agronomía, Universidad de la República - INIA
dc.relation.ispartof.es.fl_str_mv Agrociencia Uruguay. 2022;26(NE1):e504
dc.rights.license.none.fl_str_mv Licencia Creative Commons Atribución (CC - By 4.0)
dc.rights.none.fl_str_mv info:eu-repo/semantics/openAccess
dc.source.none.fl_str_mv reponame:COLIBRI
instname:Universidad de la República
instacron:Universidad de la República
dc.subject.es.fl_str_mv Capitosauroides and Pachypes-Like Tracks
Capitosauroides e Pachypes
Capitosauroides y Pachypes
Dadoxylon-Like Wood
Estratigrafia da Formação Yaguarí
Estratigrafía de Formación Yaguarí
Guadalupian-Lopingian Boundary
Huellas Karoopes
Karoopes
Limite Guadalupiano-Lopingiano
Límite Guadalupiano-Lopingiano
Madeiras Dadoxylon
Maderas Dadoxylon
Paleoambientes
Paleoenvironments
Pegadas Karoopes
Uruguai
Uruguay
Yaguarí Formation Stratigraphy
dc.title.none.fl_str_mv Enigmatic wood and first evidence of tetrapods in the Yaguarí Formation (Middle-Late Permian), Uruguay
Madera enigmática y primera evidencia de tetrápodos en Formación Yaguarí (Pérmico Medio-Superior), Uruguay
Madeira enigmática e primeira evidência de tetrápodes na Formação Yaguarí (Permiano Médio-Superior), Uruguai
dc.type.es.fl_str_mv Artículo
dc.type.none.fl_str_mv info:eu-repo/semantics/article
dc.type.version.none.fl_str_mv info:eu-repo/semantics/publishedVersion
description This article describes an intriguing fossil material recently found in the fine-grained sandstone levels from the upper section of the Yaguarí Formation of Uruguay. The nature of this specimen and its relationships were, initially, not easy to determine with confidence although it was subjected to several analyses, such as morphological and comparative examination complemented with preparation of thin sections and microanatomical and chemical studies under SEM. However, the anatomical structure shown in the performed transversal thin sections allowed solving the issue on the different animal or plant interpretations of the fossil specimen, which was finally discerned in favor of the latter. Therefore, after a detailed comparative study of the thin sections, we concluded that the specimen is a wood fragment related to the Dadoxylon-Araucarioxylon complex. Moreover, in order to provide a better biostratigraphic calibration for these deposits until now devoid of fossils, we include a preliminary description of recently found fossil footprints, which represent the first evidence of the presence of tetrapods in levels of the Yaguarí Formation (sensu Elizalde et al., 1970), below the conglomerates that have yielded the Colonia Orozco Fauna. The footprints were preliminary assigned to the ichnotaxa cf. Karoopes isp, cf. Capitosauroides isp., and cf. Pachypesisp., suggesting the presence of gorgonopsids and therocephalian therapsids and pareiasauromorphs, respectively. The Middle-Late Permian age recently suggested for the Yaguarí Formation based on paleomagnetic and radioisotopic studies is here supported by the described fossils, which thus constitute preliminary but nonetheless relevant discoveries for deposits whose fossiliferous potential has yet to be explored.
eu_rights_str_mv openAccess
format article
id COLIBRI_2ad5107cf7f0770369269df52318cdbe
identifier_str_mv Piñeiro, G. H., Marchetti, L., Marmol, S., y otros. Enigmatic wood and first evidence of tetrapods in the Yaguarí Formation (Middle-Late Permian), Uruguay. Agrociencia Uruguay, 26(NE1), e504. https://doi.org/10.31285/AGRO.26.504
2730-5066
10.31285/AGRO.26.504
instacron_str Universidad de la República
institution Universidad de la República
instname_str Universidad de la República
language eng
language_invalid_str_mv en
network_acronym_str COLIBRI
network_name_str COLIBRI
oai_identifier_str oai:colibri.udelar.edu.uy:20.500.12008/43673
publishDate 2022
reponame_str COLIBRI
repository.mail.fl_str_mv mabel.seroubian@seciu.edu.uy
repository.name.fl_str_mv COLIBRI - Universidad de la República
repository_id_str 4771
rights_invalid_str_mv Licencia Creative Commons Atribución (CC - By 4.0)
spelling Piñeiro Graciela Helena, Universidad de la República, Facultad de Ciencias, Departamento de Paleontología, Montevideo, Uruguay https://orcid.org/0000-0002-5872-9721Marchetti Lorenzo, Museum für Naturkunde Berlin, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Berlin, Germany https://orcid.org/0000-0002-1047-7887Marmol Sebastián, Universidad de la República, Facultad de Agronomía, Departamento de suelos y aguas, Montevideo, Uruguay https://orcid.org/0000-0001-7059-2724Celio Antonella, Universidad de la República, Facultad de Agronomía, Departamento de suelos y aguas, Montevideo, Uruguay https://orcid.org/0000-0002-1425-0217Xavier Pedro Luis, Universidade Federal do Rio Grande do Sul, Programa de Pós-Graduação em Geociências, Porto Alegre, Brasil https://orcid.org/0000-0001-9090-5361Francia Martin, Universidad de la República, Facultad de Agronomía, Departamento de suelos y aguas, Montevideo, Uruguay https://orcid.org/0000-0001-6107-2815Schultz Cesar Leandro, Universidade Federal do Rio Grande do Sul, Departamento de Estratigrafia e Paleontologia, Porto Alegre, Brasil https://orcid.org/0000-0001-7121-04092024-04-29T14:56:21Z2024-04-29T14:56:21Z2022Piñeiro, G. H., Marchetti, L., Marmol, S., y otros. Enigmatic wood and first evidence of tetrapods in the Yaguarí Formation (Middle-Late Permian), Uruguay. Agrociencia Uruguay, 26(NE1), e504. https://doi.org/10.31285/AGRO.26.5042730-5066https://hdl.handle.net/20.500.12008/4367310.31285/AGRO.26.504This article describes an intriguing fossil material recently found in the fine-grained sandstone levels from the upper section of the Yaguarí Formation of Uruguay. The nature of this specimen and its relationships were, initially, not easy to determine with confidence although it was subjected to several analyses, such as morphological and comparative examination complemented with preparation of thin sections and microanatomical and chemical studies under SEM. However, the anatomical structure shown in the performed transversal thin sections allowed solving the issue on the different animal or plant interpretations of the fossil specimen, which was finally discerned in favor of the latter. Therefore, after a detailed comparative study of the thin sections, we concluded that the specimen is a wood fragment related to the Dadoxylon-Araucarioxylon complex. Moreover, in order to provide a better biostratigraphic calibration for these deposits until now devoid of fossils, we include a preliminary description of recently found fossil footprints, which represent the first evidence of the presence of tetrapods in levels of the Yaguarí Formation (sensu Elizalde et al., 1970), below the conglomerates that have yielded the Colonia Orozco Fauna. The footprints were preliminary assigned to the ichnotaxa cf. Karoopes isp, cf. Capitosauroides isp., and cf. Pachypesisp., suggesting the presence of gorgonopsids and therocephalian therapsids and pareiasauromorphs, respectively. The Middle-Late Permian age recently suggested for the Yaguarí Formation based on paleomagnetic and radioisotopic studies is here supported by the described fossils, which thus constitute preliminary but nonetheless relevant discoveries for deposits whose fossiliferous potential has yet to be explored.Este artículo describe un intrigante fósil recientemente descubierto en niveles de areniscas finas a medias de la sección superior de la Formación Yaguarí de Uruguay. La naturaleza y las afinidades de este espécimen no fueron inicialmente fáciles de determinar, a pesar de ser sometido a numerosos estudios, incluido un análisis anatómico-comparativo complementado con la preparación de secciones delgadas y análisis de su microestructura bajo Microscopio Electrónico de Barrido. La microestructura mostrada por las láminas delgadas en corte transversal permitió finalmente resolver el problema sobre discernir entre una naturaleza animal o vegetal del espécimen, siendo la última la aceptada. El detallado estudio comparativo de las secciones delgadas reveló que el espécimen es un fragmento de madera relacionado al complejo Dadoxylon-Araucarioxylon. Asimismo, para proveer una mejor calibración temporal de los depósitos de la sección superior de la Formación Yaguarí (Miembro Villa Viñoles), también se incluye en este trabajo una descripción preliminar de huellas de tetrápodos recientemente halladas en esta unidad, las cuales representan la primera evidencia de la presencia de tetrápodos en niveles de la Formación Yaguarí (sensu Elizalde et al., 1970) subyacentes a los conglomerados portadores de la Fauna de Colonia Orozco. Las huellas fueron asignadas a los icnotaxa cf. Karoopes isp., cf. Capitosauroides isp. y cf. Pachypes isp., sugiriendo la presencia de terápsidos gorgonópsidos y therocefalidos y de pareiasauromorfos, respectivamente. La edad Pérmico Medio a Tardío recientemente sugerida para la Formación Yaguarí, basado en estudios de paleomagnetismo sustentados por dataciones radioisotópicas, es congruente con la aportada por los fósiles aquí descriptos, que, aunque fragmentarios, constituyen descubrimientos preliminarmente relevantes para depósitos cuyo potencial fosilífero no ha sido aún explorado.Este artigo descreve um intrigante material fóssil achado recentemente nos níveis de arenito fino da seção superior da Formação Yaguarí no Uruguai. A natureza deste espécime e suas relações não foram, em um primeiro momento, facilmente determináveis, apesar do mesmo ter sido submetido a várias análises. Estas incluíram avaliações preliminares comparativas e de morfologia, complementadas com preparação de lâminas delgadas, e estudos microanatômicos e químicos em MEV. A estrutura anatômica observada nas seções delgadas transversais permitiu solucionar as controversas interpretações do espécime fóssil, animal ou planta, o qual foi finalmente discernido em favor da última. Por enquanto, depois do estudo detalhado das lâminas delgadas o espécime foi relacionado a o complexo Dadoxylon-Araucarioxylon. Além disso, para permitir uma melhor calibração bioestratigráfica para estes depósitos até então desprovidos de fósseis, incluímos uma descrição preliminar de pegadas fósseis recém-encontradas, as quais são a primeira evidência a presença de tetrápodes em níveis da Formação Yaguarí (sensu Elizalde et al., 1970), abaixo dos conglomerados que forneceram a Fauna de Colonia Orozco. As pegadas foram assignadas aos ichnotaxa cf. Karoopes isp., cf. Capitosauroides isp. e cf. Pachypes isp., sugerindo a presença de terápsidos gorgonopsídeos e terocefalídeos e de pareiasauromorfos, respectivamente. Uma idade Meso-Neopermiano recentemente sugerida para a Formação Yaguarí baseada em estudos paleomagnéticos e radiométricos é aqui suportada pelos fósseis descritos, os quais, apesar de serem descobertas preliminares, fornecem dados relevantes para depósitos cujo potencial fossilífero ainda está para ser explorado.Submitted by Colombo Cabanas Stephanie (colombo@fagro.edu.uy) on 2024-04-26T12:45:27Z No. of bitstreams: 2 license_rdf: 24251 bytes, checksum: 71ed42ef0a0b648670f707320be37b90 (MD5) 2730-5066--504.pdf: 1465751 bytes, checksum: 6b62e0880b09cc79ac1c26fbd0949760 (MD5)Approved for entry into archive by Muniz Andrea (rosmeri8@hotmail.com) on 2024-04-26T14:16:30Z (GMT) No. of bitstreams: 2 license_rdf: 24251 bytes, checksum: 71ed42ef0a0b648670f707320be37b90 (MD5) 2730-5066--504.pdf: 1465751 bytes, checksum: 6b62e0880b09cc79ac1c26fbd0949760 (MD5)Made available in DSpace by Luna Fabiana (fabiana.luna@seciu.edu.uy) on 2024-04-29T14:56:21Z (GMT). No. of bitstreams: 2 license_rdf: 24251 bytes, checksum: 71ed42ef0a0b648670f707320be37b90 (MD5) 2730-5066--504.pdf: 1465751 bytes, checksum: 6b62e0880b09cc79ac1c26fbd0949760 (MD5) Previous issue date: 2022e504application/pdfenengFacultad de Agronomía, Universidad de la República - INIAAgrociencia Uruguay. 2022;26(NE1):e504Las obras depositadas en el Repositorio se rigen por la Ordenanza de los Derechos de la Propiedad Intelectual de la Universidad de la República.(Res. Nº 91 de C.D.C. de 8/III/1994 – D.O. 7/IV/1994) y por la Ordenanza del Repositorio Abierto de la Universidad de la República (Res. Nº 16 de C.D.C. de 07/10/2014)info:eu-repo/semantics/openAccessLicencia Creative Commons Atribución (CC - By 4.0)Capitosauroides and Pachypes-Like TracksCapitosauroides e PachypesCapitosauroides y PachypesDadoxylon-Like WoodEstratigrafia da Formação YaguaríEstratigrafía de Formación YaguaríGuadalupian-Lopingian BoundaryHuellas KaroopesKaroopesLimite Guadalupiano-LopingianoLímite Guadalupiano-LopingianoMadeiras DadoxylonMaderas DadoxylonPaleoambientesPaleoenvironmentsPegadas KaroopesUruguaiUruguayYaguarí Formation StratigraphyEnigmatic wood and first evidence of tetrapods in the Yaguarí Formation (Middle-Late Permian), UruguayMadera enigmática y primera evidencia de tetrápodos en Formación Yaguarí (Pérmico Medio-Superior), UruguayMadeira enigmática e primeira evidência de tetrápodes na Formação Yaguarí (Permiano Médio-Superior), UruguaiArtículoinfo:eu-repo/semantics/articleinfo:eu-repo/semantics/publishedVersionreponame:COLIBRIinstname:Universidad de la Repúblicainstacron:Universidad de la RepúblicaPiñeiro, Graciela HelenaMarchetti, LorenzoMarmol, SebastiánCelio, AntonellaXavier, Pedro LuisFrancia, MartinSchultz, Cesar LeandroPérez-Bidegain, MarioGaucher, ClaudioLICENSElicense.txtlicense.txttext/plain; charset=utf-84267http://localhost:8080/xmlui/bitstream/20.500.12008/43673/5/license.txt6429389a7df7277b72b7924fdc7d47a9MD55CC-LICENSElicense_urllicense_urltext/plain; charset=utf-844http://localhost:8080/xmlui/bitstream/20.500.12008/43673/2/license_urla0ebbeafb9d2ec7cbb19d7137ebc392cMD52license_textlicense_texttext/html; charset=utf-820438http://localhost:8080/xmlui/bitstream/20.500.12008/43673/3/license_text3fd1ebed2dc74f31b189736d0c5b4212MD53license_rdflicense_rdfapplication/rdf+xml; charset=utf-824251http://localhost:8080/xmlui/bitstream/20.500.12008/43673/4/license_rdf71ed42ef0a0b648670f707320be37b90MD54ORIGINAL2730-5066--504.pdf2730-5066--504.pdfapplication/pdf1465751http://localhost:8080/xmlui/bitstream/20.500.12008/43673/1/2730-5066--504.pdf6b62e0880b09cc79ac1c26fbd0949760MD5120.500.12008/436732024-04-29 11:56:21.673oai:colibri.udelar.edu.uy:20.500.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Universidadhttps://udelar.edu.uy/https://www.colibri.udelar.edu.uy/oai/requestmabel.seroubian@seciu.edu.uyUruguayopendoar:47712024-07-25T14:47:58.043772COLIBRI - Universidad de la Repúblicafalse
spellingShingle Enigmatic wood and first evidence of tetrapods in the Yaguarí Formation (Middle-Late Permian), Uruguay
Piñeiro, Graciela Helena
Capitosauroides and Pachypes-Like Tracks
Capitosauroides e Pachypes
Capitosauroides y Pachypes
Dadoxylon-Like Wood
Estratigrafia da Formação Yaguarí
Estratigrafía de Formación Yaguarí
Guadalupian-Lopingian Boundary
Huellas Karoopes
Karoopes
Limite Guadalupiano-Lopingiano
Límite Guadalupiano-Lopingiano
Madeiras Dadoxylon
Maderas Dadoxylon
Paleoambientes
Paleoenvironments
Pegadas Karoopes
Uruguai
Uruguay
Yaguarí Formation Stratigraphy
status_str publishedVersion
title Enigmatic wood and first evidence of tetrapods in the Yaguarí Formation (Middle-Late Permian), Uruguay
title_full Enigmatic wood and first evidence of tetrapods in the Yaguarí Formation (Middle-Late Permian), Uruguay
title_fullStr Enigmatic wood and first evidence of tetrapods in the Yaguarí Formation (Middle-Late Permian), Uruguay
title_full_unstemmed Enigmatic wood and first evidence of tetrapods in the Yaguarí Formation (Middle-Late Permian), Uruguay
title_short Enigmatic wood and first evidence of tetrapods in the Yaguarí Formation (Middle-Late Permian), Uruguay
title_sort Enigmatic wood and first evidence of tetrapods in the Yaguarí Formation (Middle-Late Permian), Uruguay
topic Capitosauroides and Pachypes-Like Tracks
Capitosauroides e Pachypes
Capitosauroides y Pachypes
Dadoxylon-Like Wood
Estratigrafia da Formação Yaguarí
Estratigrafía de Formación Yaguarí
Guadalupian-Lopingian Boundary
Huellas Karoopes
Karoopes
Limite Guadalupiano-Lopingiano
Límite Guadalupiano-Lopingiano
Madeiras Dadoxylon
Maderas Dadoxylon
Paleoambientes
Paleoenvironments
Pegadas Karoopes
Uruguai
Uruguay
Yaguarí Formation Stratigraphy
url https://hdl.handle.net/20.500.12008/43673